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In case of O 2 , PQQH 2 acts as a catalyst in O 2 -quenching processes and quickly transform -tocopheroxyl radicals into -tocopherol (Ouchi et al., 2013) by mitigating pro-oxidant effects of -tocopheroxyl radicals (Liu et al., 2021)

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While significant breakthroughs have been achieved in multimodal modulation of mitochondrial biogenesis, mitophagy, and dynamics, persisting challenges include incomplete elucidation of molecular regulatory networks and genetic foundations, limitations of single-pathway intervention strategies in achieving multi-target coordination, and technical barriers imposed by the selective permeability of the blood-brain barrier (BBB) that hinder clinical translation of mitochondrial-targeted therapies (Meng et al., 2024)

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Ice crystal formation during freezing mechanically disrupts the liquid matrix

Several compounds have been shown to interfere with melanosome transport or to activate autophagy in cultured melanocytes and pigmented skin models (Supplementary information 4.1)