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The first group, which includes vildagliptin and saxagliptin, forms covalent bonds with both the S1 subsite (where Ser630 is located) and the S2 subsite

(3) Condensation of the tail and head group: Once both the benzoquinone head group precursor (4-HB) and the isoprenoid side chain (polyprenyl pyrophosphate, PPP) are synthesized, the enzyme COQ2 catalyzes their condensation via an electrophilic aromatic substitution 33

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