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The transcription regulator, forkhead box class O isoforms 4 (FOXO4), maintains the viability of senescent cells by localizing adjacent to p53-pS15 to prevent the nuclear exclusion of active p53-pS15 to mediate apoptosis 11

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The N-terminal domain of the target-binding site has two residues, Glu 5 and Glu 9, that are the most highly conserved in S100 proteins (136)