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Database searching was performed in Proteome Discoverer (PD, Thermo Fisher Scientific) v3.0 against the UniProt human reference proteome, allowing tryptic peptides (730 aa), up to two missed cleavages, and up to three variable modifications per peptide

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*Correspondence: Li-min Yue, [email protected] Nie, [email protected] This article was submitted to Reproduction, a section of the journal Frontiers in Physiology Disclaimer All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers

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